There are only two visual paths, translatable in greater simplicity and clarity. The centered design is more permissive, the lack of conditioning regarding folder-subfolder navigation and not having to align each quadrant's content to the left makes it more agile. It's difficult to even imagine the effort that a user with an adapted mouse must make to follow the path of this diagram. But when it comes to something more global such as a work application or even more an operating system, the design must meet optimal accessibility standards and tested to the maximum. When we analyze an interface we tend to think of "conventional" users who use a mouse, touch-pad or fingers. The user is pushed to follow a type of navigation from left to right, from bottom to top and then from top to bottom in each quadrant, scrolling down to see the content without knowing where it ends, a real chaos: 
System that sometimes works and sometimes does not, and the design of the start content is a clear example. I always had the feeling that Windows has a hard time leaving the navigation system based on the stagged of folders and subfolders from left to right. In the same way the question refers to the lack of justification for the relocation of the centered start button, I have personally always questioned the layout of the content located on the left. Well, beyond the fact that there's a better adaptability for devices with a touch-pad as mentioned in the question, I see other specifications regarding design.
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